One of the most famous autonomous differential equations is the logistic model, which describes population growth under limited resources. The equation is:
where \(P\) is the population at time \(t\text{,}\)\(r\) is the growth rate, and \(K\) is the carrying capacity, the maximum population that the environment can sustain.
Populations grow when between \(0\) and \(K\text{,}\) and shrink when above \(K\text{.}\) The model predicts eventual stabilization at the carrying capacity.
where \(N\) is the carrying capacity and \(k\) is the growth-rate constant. This form emphasizes that growth is proportional to both the current population and the remaining resource capacity \((1 - P/N)\text{.}\)
This is still a first-order autonomous differential equation, but unlike the exponential model, it is nonlinear since the right-hand side is not a linear function of \(P\text{.}\)
Let us now consider a variation of the logistic model that incorporates an important ecological detail: what happens to very small populations. The standard logistic model assumes that populations always grow when small, but in reality, sparse populations can have difficulty finding mates, making growth negative at low levels.
As an example, consider pine squirrels in the Rocky Mountains. These animals are territorial, and both overcrowding and underpopulation can limit their growth:
If the population is too large, territorial conflict slows growth or even causes decline.